Abstract
Two density experiments using Deli x AVROS and Deli x Ghana were planted
in 1990 at two localities in Costa Rica: Coto (South Pacific) and Quepos
(Central Pacific, drier area with higher solar radiation than in Coto).
In both sites, density treatments were arranged in a 'fan' pattern
ranging from 98 to 231 palms/ha. Vegetative growth and yield components
were recorded. Annual average yield (bunch weight) per palm and yield (FFB)
per hectare was higher in DxA in Coto, but not in Quepos; where DxG
performed better. The highest annual yields per palm were obtained
within a range of densities between 70 and 139 palms/ha during the first
eight years of production. The best yields (FFB) per hectare (average of
years 5 to 10) were obtained with 139 palms/ha in DxA and 164 palms in
DxG.
The first growth variable that seemed to be affected by interplant
competition with increasing plant density was leaf length (increasing),
later, a reduction in leaf emission rate was observed (around the
seventh year). Stem height was affected only in adult palms (10 or more
years).
Introduction
V ariable density experiments in oil palm were proposed by Goh (1977),
where the use of plots of different size with the same number of plants
has the advantage of a reduction in land and costs when compared with
block designs. The main disadvantage of this arrangement of plots is
that it is systematic, so no regular variance analysis can be conducted.
However, this author also described an analysis procedure to select,
between several regression equations, the best density related with
maximum yield (Goh, 1982). This analysis was then complemented with a
graphical interpretation of the results in order to select the treatment
with maximum total yields.
Smith, Donough and Corley (1996) used similar procedures to interpret
the results of a density experiment in clones with different canopy
characteristics. They found that rachis length and leaf area index could
be used as indicators for optimum density. Donough and Kwan (1991)
compared the performance of several varieties planted at different
densities in two localities, and found an interaction site by variety.
They also suggested that total oil produced per area could be a better
indicator of the effects of planting density, since fruit to bunch ratio
increased with density, which conducted to an increase in the amount of
oil produced per unit area within certain range of densities. Also
Breure (1982) showed, in a density x fertilizer trial, that rachis
length and leaf production were affected by planting density five years
after planting.
This paper compares the performance of two commercial oil palm varieties
(Deli x AVROS and Deli x Ghana) planted at different densities
(systematic fan arrangement) in two localities with different
environmental conditions (South and Central Pacific of Costa Rica).
Materials and Methods
Two density experiments using the varieties Deli x AVROS (DxA) and Deli
x Ghana (DxG) were planted in Costa Rica in June 1990 in Coto (South
Pacific) and Quepos (Central Pacific coast). In both cases, the
densities evaluated covered a range from 98 to 231 palms per hectare;
even though the total density range varied from 70 to 326 palms per
hectare, but these extreme densities were used as border plots. In Coto,
there were three six-palm plots for each variety (total of 18 palms for
each density and variety). In Quepos, four plots were planted for each
variety (24 palms for each variety and density).
The mean annual rainfall in Coto is 3,930 mm, with only two months with
rainfall below 100 mm. Under these conditions there is no water deficit
or it is very mild in some years. Mean monthly temperatures are within a
suitable range for oil palm cultivation (minimum 20-22 °C and maximum
31-34 °C). In this locality solar radiation is below optimal range (<15
Mj/m˛) for 4 to 6 months. In Quepos, total annual rainfall is similar
(3,674 mm), but there is a dry season from December to April, with a
total water deficit of around 400 mm. Mean monthly temperatures vary
from a minimum of 23 °C to a maximum of 31 °C.
Yield was recorded during nine (Coto) and eight years (Quepos).
Vegetative growth (leaf length and trunk height) was recorded at 76 (Quepos)
and 80 (Coto) months after planting to compare both localities; but in
Coto growth was also evaluated at 35, 83 and 133 months after planting.
Bunch components were determined from about 50 bunches from each density
in Coto, when palms were ten years old.
Results and discussion
Yield
The average annual yield per palm for three density ranges was used to
facilitate the interpretation of the results: 1) 'low' to 'normal'
densities were those varying from 98 to 164 palms/ha, 2) 'high
densities' were those varying from 164 to 231 palms/ha, and 3) 'the
total range of densities' covered from 98 to 231 palms/ha
(Table. 1).
DxA, a fast growing variety, performed better at low densities when
planted in the Coto locality (less solar radiation). The average annual
yield per palm in Coto was higher in Deli x AVROS (124.5 kg FFB) when
compared with Deli x Ghana (103.7 kg). However, a different situation
was observed in Quepos (drier area with higher solar radiation), where
annual yield per palm in Ghana (129.7 kg) was higher compared with AVROS
(119.4 kg). Yield per palm for DxA was similar in both localities, but
DxG performed better in Quepos, indicating, perhaps, its higher
tolerance to water deficit.
Figure 1
shows the reduction in yield per
palm as plant density increased.
Optimum density for best yields according to palm age. This was
estimated by determining the maximum yield (per palm and per hectare)
using mobile means for three years. During the first ten years, the
maximum yield per palm was recorded at lower densities in Coto when
compared with Quepos (70 to 83 palms/ha in Coto vs. 98 palms/ha in
Quepos), which was expected for the most vigorous growth of DxA
(Table 2).
Best average FFB yields/ha/year (palms from five to 10 years old) were
obtained in Quepos at higher densities than in Coto (164 to 231 palms/ha
in Quepos vs. 139 to 164 in Coto). In both locations, the optimum yield
for DxG was obtained at the highest densities (Table 2).
It was apparent that both varieties could have been planted at higher
densities in Quepos in order to obtain the highest yields per hectare.
Considering this, the optimum densities to reach the best yields in Coto
were 139 palms/ha for DxA and 164 for DxG; which agreed with the present
knowledge on the behavior of these two varieties (Breure, 2007).
However, other factors besides palm density could have also affected
yields (soil types, fertilization practices, rainfall, solar radiation…)
but they were not considered in the present analysis.
Vegetative growth and bunch composition
When palms were about six years of age, leaf length was shorter in palms
planted at the lower densities, but trunk height had not been apparently
affected yet. The difference in leaf length between DxA and DxG was
larger in Coto (108 cm) than in Quepos (20 cm). A faster trunk growth of
DxA in Coto was expected considering a much lower solar radiation in
this locality. However, in Quepos, water deficit seemed to had adversely
affected growth in DxA, and no clear differences were noted between the
two varieties
(Table 3).
Leaf emission rate was higher in Quepos (28-29 leaves/year vs. 21-24 in
Coto). The magnitude of this variable decreased as plant density
increased in Coto, but not in Quepos
(Table 4).
As seen in Coto, trunk height became an indicator of interplant
competition only when palms reached maturity
(Fig. 2a and
Fig. 2b). However,
leaf length showed clear signs of light competition even in rather young
palms when planted at higher densities. For the fast growing variety DxA,
leaf length was 13-21 % shorter in palms planted at 98 palms/hectare,
when compared to the density of 231 palms/ha. For DxG, light competition
was less evident (leaves 10 to 16% shorter)
(Table 5,
Fig. 3a and
Fig. 3b).
The variety Deli x Ghana showed a higher oil to bunch ratio, which was
independent of plant density. Fuit set increased with palm density
(Table 6
), which could indicate a more favorable environment for
pollination as suggested by Donough and Kwan (1991). Breure et al (1990)
also indicated that the pollinating weevils were more active in the cool
and shaded conditions prevalent at higher densities.
Conclusions
The variety Deli x Ghana showed some important advantages over the
traditional variety Deli x AVROS, especially more tolerance to light
competition (can be planted at higher densities due to its shorter
leaves as seen in Coto), tolerance to drought (performed better in an
area with high water deficit) and has a higher oil content.
Best economical yields could be obtained over a range of planting
densities; but, for a vigorous variety like DxA, best yields were
obtained at lower densities when planted in a site like the South
Pacific coast of Costa Rica were solar radiation could be a limiting
factor. The standard density of 142 or less palms per hectare can be
considered acceptable for DxA in this environment. For a less vigorous
variety like DxG, with shorter leaves, planting density can be increased
to 160 plants per hectare.
The first vegetative variable that seemed to be affected by interplant
competition with increasing plant density was leaf length (increased).
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